There is a difference writing for the web from print, when you write headlines on the web one of the most important thing about writing a headline is that it includes key words so that it is easy to search. Things like alliteration, puns, cryptic, a play on words but you cannot do that writing for the web. You should make it direct and straightforward when online so that it is easy to search online. When writing headlines you not need punctuation, no full stops. When you are quoting someone only use single quote marks, you can use a comma. For capital letters, traditionally cap up the words.
Newspapers & their headlines:
The Telegraph caps just at the beginning
The Sun all caps
The Guardian only caps at the beginning
Daily Mail only caps at the beginning
Graduate multimedia journalist and contributing writer at Culturefly